Aki's twin sister Aya is the reincarnation of Ceres, and she is eventually led to killing him to save both herself and Ceres and release Aki from Shisou's control.
His father's crazy sister Augusta is.
And then they learn that they aren't blood-related after all--in a very mind-twisting way.
In fact, when her actual older brother Tokio comes to take her in, it's right when ; she then uses this to get herself free, call out her neglectful adoptive parents, and leave the foster family.
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No, they're full-blown siblings.